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Cyperaceae

Perennial or annual. Stems bearing inflorescences usually solid and triangular in transverse section, rarely terete or jointed. Leaves usually in 3 rows, mostly basal, either scale- or sheathlike, or linear and grass-like, mostly withouth a ligule. Inflorescence variable; from a terminal few-flowered spikelet to a compound panicle with many spikelets, either naked or with leaf-like involucral bracts. Spikelets unisexual or bisexual, few- to many-flowered, often with many sterile flowers. Flowers unisexual or bisexual, solitary within a flume-like bract. Perianth reduced to hairs, bristles, scales or absent, sometimes forming a sac (utricle) surrounding the nut. Stamens usually 3, rarely 1-12 or more. Ovary superiour, with 1 erect ovule; style 1; stigmas 2 or 3; fruit a nut, either trigonous or biconvex, often with a persistent beak.

There are many different interpretations of the Cyperoid flower. In this account the flowers are considered to be basically bisexual with a perianth reduced to scales or bristles; the number of stamens in the flower is considered to be 3, in some cases reduced to one; many stamens arise by the grouping of a number of one-stamened flowers. Generic delimitation is problematic, the treatment adopted here takes Cyperus in its broadest sense, and Scirpus in a narrow sense (Eleogiton, Isolepis, Oxycaryum, Schoennoplectus and Websteria are recognised as genera). Unless stated the genera are assumed to be helophyted, pollinated by wind (anemophilous), having nuts as diaspores with unknown dispersal mechanisms.

© Christopher D.K. Cook