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Ceratophyllaceae

Entirely submerged perennial or sometimes annual. Roots absent (even in the embryo) but branches sometimes modified as 'rhizoids'. Stem branched but not more than one branch at a node. Leaves whorled, (3-) 6-8 (-10) at a node, rather rigid, often brittle, 1-4 times forked, with 2 rows of minute teeth along the ultimate segments, each segment tipped by 2 bristles. Flowers unisexual, solitary in the axil of 1 leaf in a whorl. Perianth of 8-12 linear segments, united at the base. Stamens numerous; filaments short or absent; connective prolonged at the apex. Ovary superior, 1-locular; fruit a 1-seeded nut, tipped by the persistent style and often with additional basal and marginal spines, sometime also warty.

Totally submerged, rootless, free-swimming: hypohydrophilous, stamens are liberated and dehisce floating just below the water surface: diaspores relatively large and spiny nuts or specialized buds, dispersal probably by animals and by water: large populations may be troublesome blocking or constricting water channals.

© Christopher D.K. Cook